When you set an load the gun , you have one round in the barrel ( first shot ) and one under the extractor ( 2nd shot ) . The gun loads the third and after shots.
Your hand working the bolt with no problems is supporting this , the gun itself is not feeding itself correctly.
Is the first gun round ( third shot ) under the extractor or no ? If under , belt is feeding over , if not , belt isn't ( feed pawls spring too strong or feed pawl arm not moving right . Some guns just perfer which way the lins go . If using belt , do you have a trunion protector ? Combo may be too tight .)
If under extractor , could be worn/weak extractor spring or feedway spacer is not compatable with the ammo ( over all length allowing ammo groove to move too far forward to be caught by extractor ) . The greater speed of live fire may not allow a weak spring to grab .
Chris